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Once in a generation, a Wallace may be found physically, mentally, and morally qualified to wander unscathed through the tropical wilds of America and of Asia, to form magnificent collections as he wanders, and withal to think out sagaciously the conclusions suggested by his collections; but, to the ordinary explorer or collector, the dense forests of equatorial Asia and Africa, which constitute the favorite habitation of the Orang, the Chimpanzee, and the Gorilla, present difficulties of no ordinary magnitude; and the man who risks his life by even a short visit to the malarious shores of those regions may well be excused if he shrinks from facing the dangers of the interior; if he contents himself with stimulating the industry of the better-seasoned natives, and collecting and collating the more or less mythical reports and traditions with which they are too ready to supply him.

The fact of coadaptation, which was supposed to furnish the strongest argument against the principle of selection, in reality yields the clearest evidence in favour of it. We _must_ assume it, _because no other possibility of explanation is open to us, and because these adaptations actually exist, that is to say, have really taken place_. With this conviction I attempted, as far back as 1894, when the idea of germinal selection had not yet occurred to me, to make "harmonious adaptation" (coadaptation) more easily intelligible in some way or other, and so I was led to the idea, which was subsequently expounded in detail by Baldwin, and Lloyd Morgan, and also by Osborn, and Gulick as _Organic Selection_. It seemed to me that it was not necessary that all the germinal variations required for secondary variations should have occurred _simultaneously_, since, for instance, in the case of the stag, the bones, muscles, sinews, and nerves would be incited by the increasing heaviness of the antlers to greater activity in _the individual life_, and so would be strengthened. The antlers can only have increased in size by very slow degrees, so that the muscles and bones may have been able to keep pace with their growth in the individual life, until the requisite germinal variations presented themselves. In this way a disharmony between the increasing weight of the antlers and the parts which support and move them would be avoided, since time would be given for the appropriate germinal variations to occur, and so to set agoing the _hereditary_ variation of the muscles, sinews and bones.[42]

The leaders of the popular party perceived the mistake they had made in alienating the Italians from their cause, and they now secured their adhesion by offering them the Roman citizenship if they would support the Agrarian Law. As Roman citizens they would, of course, be entitled to the benefits of the law, while they would, at the same time, obtain what they had so long desired--an equal share in political power. But the existing citizens, who saw that their own importance would be diminished by an increase in their numbers, viewed such a proposal with the utmost repugnance. So strong was their feeling that, when great numbers of the Italians had flocked to Rome in B.C. 126, the Tribune M. Junius Pennus carried a law that all aliens should quit the city.

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