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The Navaho are a pastoral, semi-nomadic people whose activities centre in their flocks and small farms. Their reservation of more than fourteen thousand square miles is the desert plateau region of northern Arizona and New Mexico. Its mesas and low mountains are sparsely covered with pinon and cedar, and on the higher levels are small but beautiful forests of pine. Back and forth in all parts of this vast region the Navaho drive their flocks. At the season when the slight rainfall produces even scant pasturage on the desert plains the flocks are pastured there; but as the grass becomes seared by the summer sun and exhausted from pasturing, the flocks are taken into the mountains, where the shade of the pines lends grateful coolness. Again, as the deep snows of winter come, the sheep and goats are driven down to the wooded mesas, where there is little snow and an abundance of fuel, of which there is none on the plains. And so, year in, year out, the flocks slowly drift back and forth from plain to mesa and from mesa to mountain.

It is not surprising under these circumstances to learn that sticklebacks are successful and increasing animals. Their numbers are enormous, wherever they get a fair chance in life, because they multiply rapidly up to the extreme limit of the means of subsistence, and develop as fast as food remains for them. There the inexorable Malthusian law at last steps in: when there is not food enough for all some must starve; that is the long and the short of the great population question. But while provender is forthcoming they increase gayly. Sticklebacks live mainly on the spawn of other fish, though they are so careful of their own, and they are therefore naturally hated by trout-preservers and owners of fisheries in general. Thousands and thousands are caught each year; in some places, indeed, they are so numerous that they are used as manure. It is their numbers, of course, that make them formidable; they are the locusts of the streams, well armed and pugnacious, and provided with most remarkable parental instincts of a protective character, which enables them to fill up all vacancies in their ranks as fast as they occur with astonishing promptitude.

Although the ant is a small animal as compared with man or with the Irish Elk, the "soldier" with its relatively enormous jaws is hardly less heavily burdened than the Elk with its antlers, and in the ant's case, too, a strengthening of the skeleton, of the muscles, the nerves of the head, and of the legs must have taken place parallel with the enlargement of the jaws. _Harmonious adaptation_ (coadaptation) has here been active in a high degree, and yet these "soldiers" are sterile! There thus remains nothing for it but to refer all their adaptations, positive and negative alike, to processes of selection which have taken place in the rudiments of the workers within the egg and sperm-cells of their parents. There is no way out of the difficulty except the one Darwin pointed out. He himself did not find the solution of the riddle at once. At first he believed that the case of the workers among social insects presented "the most serious special difficulty" in the way of his theory of natural selection; and it was only after it had become clear to him that it was not the sterile insects themselves but their parents that were selected, according as they produced more or less well adapted workers, that he was able to refer to this very case of the conditions among ants "_in order to show the power of natural selection_."[40] He explains his view by a simple but interesting illustration. Gardeners have produced, by means of long continued artificial selection, a variety of Stock, which bears entirely double, and therefore infertile flowers.[41] Nevertheless the variety continues to be reproduced from seed, because, in addition to the double and infertile flowers, the seeds always produce a certain number of single, fertile blossoms, and these are used to reproduce the double variety. These single and fertile plants correspond "to the males and females of an ant-colony, the infertile plants, which are regularly produced in large numbers, to the neuter workers of the colony."

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